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I wonder if its to do with the fact these females were all born where they live now, so have had the same management(and female companions) throughout their life-no stress or disturbance from being moved to other locations and they still live in a herd situation, although its less than half the size it used to be.
Many other parks have even smaller numbers and the zoo-bred offspring have possibly been moved about a lot more. Are there stress/incompatability factors at work in the zoobred offspring?
Maybe, but then wouldn't the moving of wild born animals be potential more stressful? Were as zoo bred animals are a little more use to louder noises. Therefore more likely not to get as stressed being moved about?