The Zoochat Photographic Guide to Rodents: part two

This thread is a continuation of The Zoochat Photographic Guide to Rodents: part one


There are approximately 2400 species in the Order Rodentia, divided between five Suborders. The first four Suborders - Sciuromorpha, Anomaluromorpha, Castorimorpha, and Hystricomorpha - were covered in the previous thread. The final Suborder, the subject of this thread, is Myomorpha which contains about 1680 species.


Part one:
Suborder Sciuromorpha (c.330 spp)
Suborder Anomaluromorpha (9 spp)
Suborder Castorimorpha (c.100 spp)
Suborder Hystricomorpha (c.300 spp)

Part two:
Suborder Myomorpha (c.1680 spp)


There was a fairly good representation of photographs for part one - over half the genera and about a quarter of the individual species were represented by photographs - but part two is much less favourable as it is largely composed of rats and mice which are rarely kept in zoos and rarely photographed in the wild.


For the most part I will probably forego the inclusion of subspecies due to the sheer number of species and the taxonomic complexities. For those species which are represented by photographs I will endeavour to sort out the subspecies, but for the un-depicted species I will mostly leave them as simple lists.

 

The presentation of Families as they appear in these two threads are as follows.


Suborder SCIUROMORPHA (c.330 spp)
Family Aplodontiidae - Sewellel or Mountain Beaver (1 sp)
Family Sciuridae - Squirrels (c.300 spp)
Family Gliridae - Dormice (c.30 spp)


Suborder ANOMALUROMORPHA (9 spp)
Family Anomaluridae - Anomalures or Scaly-tailed Squirrels (7 spp)
Family Pedetidae - Springhares (2 spp)


Suborder CASTORIMORPHA (c.100 spp)
Family Castoridae - Beavers (2 spp)
Family Geomyidae - Pocket Gophers (c.35-40 spp)
Family Heteromyidae - Kangaroo Rats and Kangaroo Mice (c.60 spp)


Suborder HYSTRICOMORPHA (c.300 spp)
Family Diatomyidae / Laonastidae - Laotian Rock Rat (1 sp)
Family Ctenodactylidae - Gundis (5 spp)
Family Bathyergidae - Blesmols (c.25 spp)
Family Hystricidae - Old World Porcupines (11 spp)
Family Petromuridae - Dassie Rat (1 sp)
Family Thryonomyidae - Cane Rats (2 spp)
Family Erethizontidae - New World Porcupines (c.17 spp)
Family Chinchillidae - Chinchillas and Viscachas (7 spp)
Family Dinomyidae - Pacarana (1 sp)
Family Caviidae - Cavies (c.20 spp)
Family Dasyproctidae - Agoutis (c.13 spp)
Family Cuniculidae - Pacas (3 spp)
Family Ctenomyidae - Tuco-tucos (c.65 spp)
Family Octodontidae - Octodonts (14 spp)
Family Abrocomidae - Chinchilla Rats (c.10 spp)
Family Echimyidae - Spiny Rats (c.90 spp)
Family Myocastoridae - Coypu (1 sp)
Family Capromyidae - Hutias (c.13 spp)


Suborder MYOMORPHA (c.1680 spp)
Family Dipodidae - Jumping Mice and Jerboas (c.50 spp)
Family Platacanthomyidae - Spiny Dormice (3 spp)
Family Spalacidae - Mole-Rats and Bamboo Rats (c.30-40 spp)
Family Calomyscidae - Mouse-like Hamsters (8 spp)
Family Nesomyidae - Madagascan and African Rats and Mice (c.65-70 spp)
Family Cricetidae - Voles, Hamsters, New World Rats and Mice (c.700 spp)
Family Muridae - Old World or "True" Mice and Rats, Gerbils, and Spiny Mice (c.810 spp)

 

Suborder MYOMORPHA


There are seven Families within this Suborder, containing about 1680 species in total. Of those, only 171 species in 91 genera are depicted in the thread.


Dipodidae (Jumping Mice and Jerboas)
About 50 species in 16 genera, of which 13 species from 8 genera are depicted here.


Platacanthomyidae (Spiny Dormice)
Three species in two genera, of which one species is depicted here.


Spalacidae (Mole-Rats and Bamboo Rats)
About 30 to 40 species in six genera, of which only six species from four genera are depicted here.


Calomyscidae (Mouse-like Hamsters)
Eight species in one genus, of which one species is depicted here.


Nesomyidae (Madagascan and African Rats and Mice)
About 65 to 70 species in 21 genera, of which only seven species from six genera are depicted here.


Cricetidae (Voles, Hamsters, New World Rats and Mice)
About 700 species in about 140 genera, of which only 56 species from 33 genera are depicted here.


Muridae (Old World or "True" Mice and Rats, Gerbils, and Spiny Mice).
About 810 species in about 160 genera, of which 87 species from 38 genera are depicted here.

 

DIPODIDAE
Jumping Mice and Jerboas
About fifty species in sixteen genera, of which thirteen species from eight genera are represented in the Zoochat galleries.


The family is most commonly divided into six subfamilies, although it may also be split into two separate families: Zapodidae, with the two subfamilies Sicistinae (birch mice) and Zapodinae (jumping mice); and Dipodidae with the remaining four subfamilies (jerboas).


Sicistinae (birch mice) - about thirteen species in the single genus Sicista.
Only one species is depicted here.

Zapodinae (jumping mice) - five species in three genera (Eozapus, Napaeozapus, Zapus).
One of the species is depicted here, from the genus Zapus.

Allactaginae (four-toed and five-toed jerboas) - about sixteen species in three genera (Allactaga, Allactodipus, Pygeretmus).
There are five species depicted here, all from the genus Allactaga.

Cardiocraniinae (pigmy jerboas) - about seven species in two genera (Cardiocanius, Salpingotus).
There are two species depicted here, both from the genus Salpingotus.

Dipodinae (three-toed jerboas) - about nine species in five genera (Dipus, Eremodipus, Jaculus, Stylodipus, Paradipus).
There are four species depicted here, from three genera: Dipus, Jaculus and Stylodipus.

Euchoreutinae (Long-eared Jerboa) - one species, in the genus Euchoreutes
The single species is depicted here.


The accounts will follow the order of genera as given above (alpabetically within subfamily).

 

Sicistinae (birch mice)
About thirteen species in the single genus Sicista.

Only one species is represented in the Zoochat galleries.


Northern Birch Mouse Sicista betulina
Monotypic. Subspecies which may be included (now generally considered synonyms) include montana, norvegica, taigica, tatricus. The species Sicista strandi was formerly included within S. betulina.

Both photos by @Giant Eland at Moscow Zoo (Russia)


(Sicista betulina) Northern birch mouse - ZooChat


(Sicista betulina) Northern birch mouse - ZooChat


The remaining species in the genus are as follows (various of these may be combined or even further split by various authors):

Armenian Birch Mouse Sicista armenica
Caucasian Birch Mouse Sicista caucasica
Long-tailed Birch Mouse Sicista caudata
Chinese Birch Mouse Sicista concolor
Kazbeg Birch Mouse Sicista kazbegica
Kluchor Birch Mouse Sicista kluchorica
Altai Birch Mouse Sicista napaea
Grey Birch Mouse Sicista pseudonapaea
Severtzov's Birch Mouse Sicista severtzovi
Strand's Birch Mouse Sicista strandi
Southern Birch Mouse Sicista subtilis
Tien Shan Birch Mouse Sicista tianshanica

 

Zapodinae (jumping mice)
Five species in three genera (Eozapus, Napaeozapus, Zapus). Especially in earlier decades, the three genera have been combined under Zapus.

One of the species is represented in the Zoochat galleries (albeit only as a museum specimen).


Eozapus
One species

Chinese Jumping Mouse Eozapus setchuanus


Napaeozapus
One species

Woodland Jumping Mouse Napaeozapus insignis


Zapus
Three species


Meadow Jumping Mouse Zapus hudsonius
Eleven subspecies: acadicus, alascensis, americanus, campestris, canadensis, hudsonius, intermedius, ladas, pallidus, preblei, tenellus


Photo by @Pleistohorse at the UAF Museum of the North (USA) (taxidermy specimen, presumably of the local subspecies acadicus)


Meadow Jumping Mouse - ZooChat


The remaining two species in the genus are as follows:

Western Jumping Mouse Zapus princeps
Pacific Jumping Mouse Zapus trinotatus

 

This will more-or-less be my plan for the Eulipotyphla guide too - especially given the fact it will doubtless have an even worse ratio of photos than this thread will!

 

Allactaginae (four-toed and five-toed jerboas)
About sixteen species in three genera (Allactaga, Allactodipus, Pygeretmus).

There are five species represented in the Zoochat galleries, all from the genus Allactaga.


Allactaga
Thirteen species, five of which are depicted here.


Gobi Jerboa Allactaga bullata
Monotypic

Photo by @Deer Forest in the wild, China


Gobi jerboa - ZooChat


Small Five-toed Jerboa Allactaga elater
Up to eleven subspecies have been recognised: aralychensis, bactriana, caucasicus, dzungariae, elator, heptneri, indica, kizljaricus, strandi, turkmeni, zaisanicus. May be a species complex.

Photo by @Maguari at Plzen Zoo (Czech Republic)


Small Five-toed Jerboa at Plzen, 25/05/10 - ZooChat


Great Jerboa Allactaga major
Monotypic?

Photo by @Giant Eland at Moscow Zoo (Russia)


(Allactaga major) Great jerboa - ZooChat


Mongolian Five-toed Jerboa Allactaga sibirica
Seven to ten subspecies may be recognised: altorum, annulata, bulganensis, dementiewi, mongolica, ognevi, saltator, semideserta, sibirica, suschkini

Photo by @Deer Forest at Beijing Zoo (China)


Mongolian five-toed jerboa (Allactaga sibirica) - ZooChat


Four-toed Jerboa Allactaga tetradactyla
Monotypic

Photo by @vogelcommando in a private collection


Four-toed jerboa - ZooChat



The remaining species in this genus as are follows:

Balikun Jerboa Allactaga balikunica
Euphrates Jerboa Allactaga euphratica
Iranian Jerboa Allactaga firouzi (probably conspecific with A. hotsoni)
Hotson's Jerboa Allactaga hotsoni
Severtzov's Jerboa Allactaga severtzovi
Toussi Jerboa Allactaga toussi (probably should be included within A. elator)
Vinogradov's Jerboa Allactaga vinogradovi
Williams's Jerboa Allactaga williamsi


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Allactodipus
One species, not depicted here.

Bobrinski's Jerboa Allactodipus bobrinskii


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Pygeretmus
Three species, none of which are depicted here.

Lesser Fat-tailed Jerboa Pygeretmus platyurus
Dwarf Fat-tailed Jerboa Pygeretmus pumilio
Greater Fat-tailed Jerboa Pygeretmus shitkovi

 

Cardiocraniinae (pigmy jerboas)
About seven species in two genera (Cardiocanius, Salpingotus).

There are two species represented in the Zoochat galleries, both from the genus Salpingotus.



Cardiocranius
One species, not depicted here.

Five-toed Pigmy Jerboa Cardiocranius paradoxus



Salpingotus
Six species, two of which are depicted here.


Thick-tailed Pigmy Jerboa Salpingotus crassicauda
Two subspecies may be recognised (crassicauda, gobicus) but otherwise treated as monotypic.

Both photos by @Giant Eland at Moscow Zoo (Russia)


(Salpingotus crassicauda) thick-tailed pygmy jerboa - ZooChat


(Salpingotus crassicauda) thick-tailed pygmy jerboa - ZooChat


Kozlov's Pigmy Jerboa Salpingotus kozlovi
Two subspecies may be recognised (kozlovi, xiangi) but otherwise treated as monotypic.

Photo by @Giant Eland at Moscow Zoo (Russia)


(Salpingotus kozlovi) Kozlov's pygmy jerboa - ZooChat


The remaining four species in this genus are not represented in the Zoochat galleries:

Heptner's Pigmy Jerboa Salpingotus heptneri
Baluchistan Pigmy Jerboa Salpingotus michaelis (may be put in its own genus, Salpingotulus)
Pale Pigmy Jerboa Salpingotus pallidus
Thomas's Pigmy Jerboa Salpingotus thomasi (known only from the type specimen, and may be conspecific with S. michaelis)

 

Dipodinae (three-toed jerboas)
About nine species in five genera (Dipus, Eremodipus, Jaculus, Paradipus, Stylodipus).

There are four species represented in the Zoochat galleries, from the genera Dipus, Jaculus and Stylodipus.



Dipus
One or two species (or more)


Northern Three-toed or Hairy-footed Jerboa Dipus sagitta
IUCN says "Eight subspecies are recognized; however there is a clear geographical cline in size and colouration from north to south."
Recent papers suggest that the genus Dipus contains at least two species, D. sagitta and D. deasyi.

Photo by @Deer Forest at Beijing Zoo (China)


Northern three-toed jerboa (Dipus sagitta) - ZooChat


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Eremodipus
One species, not depicted here.


Lichtenstein's Jerboa Eremodipus lichtensteini


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Jaculus
Three species, two of which are depicted here.


Lesser Egyptian Jerboa Jaculus jaculus
Up to nineteen subspecies have been described, although there seems little agreement on treatment. At the extreme end the species is treated as being monotypic (as in Kingdon's Mammals of Africa). Some forms (e.g. deserti and schlueteri) have been considered full species.

Photo by @Mo Hassan at Zoo Plzen (Czech Republic)


Lesser Egyptian jerboa - ZooChat


Photo by @ThylacineAlive at Bronx Zoo (USA)


Lesser Egyptian Jerboa - ZooChat


Greater Egyptian Jerboa Jaculus orientalis
Monotypic

Photo by @Zooish at Singapore Zoo (Singapore)


Greater Egyptian Jerboa (Jaculus orientalis) - ZooChat



The third species, not pictured here, is the Blanford's Jerboa Jaculus blanfordi.


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Paradipus
One species, not depicted here.


Comb-toed Jerboa Paradipus ctenodactylus


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Stylodipus
Three species, one of which is depicted here.


Thick-tailed Three-toed Jerboa Stylodipus telum
Treated as monotypic by some authors, but otherwise includes the subspecies amankaragai, falzfeini, karelini, telum, turovi. Formerly treated as a subspecies was Stylodipus andrewsi.

Photo by @Chlidonias at the Melbourne Museum (Australia) (taxidermy specimen)


Tragic Taxidermy 11 - ZooChat


The other two species, not pictured here:

Andrews's Three-toed Jerboa Stylodipus andrewsi
Mongolian Three-toed Jerboa Stylodipus sungorus

 

Euchoreutinae (Long-eared Jerboa)
One species.


Long-eared Jerboa Euchoreutes naso
Monotypic

Photo by @alexkant at Moscow Zoo (Russia)


Long-eared jerboa/ Euchoreutes naso - ZooChat

 

Fixed that for you

 

PLATACANTHOMYIDAE
Spiny Dormice
At least three species in two genera, of which one species is represented in the Zoochat galleries.

Despite the common name and their general appearance, the Spiny Dormice are not related to the "true" dormice which are now placed in the Suborder Sciuromorpha alongside squirrels.


.....................................................................


Platacanthomys
One species


Malabar Spiny Dormouse Platacanthomys lasiurus
Monotypic


This species is not depicted here.


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Typhlomys
At least two species, one of which is depicted here.

Formerly just a single species was recognised (T. cinereus, with several subspecies), with the Vietnamese species T. chapensis later being split. A revision in 2017 named four species: T. chapensis, T. cinereus, T. daloushanensis, and T. nanus. In a 2021 paper, T. huangshanensis was named.


Vietnamese Pigmy Dormouse Typhlomys chapensis
Possibly two subspecies: chapensis, guangxiensis

Formerly this species was combined with the Chinese Pigmy Dormouse T. cinereus but genetic and morphological data show it to be distinct (e.g. see An enigmatic pygmy dormouse: molecular and morphological evidence for the species taxonomic status of Typhlomys chapensis (Rodentia: Platacanthomyidae)). The second subspecies listed above (guangxiensis) is most similar genetically to T. chapensis rather than T. cinereus.

The photos present in the Zoochat galleries, all taken at Moscow Zoo, are mostly labeled as being T. cinereus because they date from before the split of T. chapensis (formerly T. cinereus chapensis).


Photo by @Giant Eland at Moscow Zoo (Russia)


(Typhlomys cinereus) Chinese pygmy dormouse - ZooChat

Photo by @alexkant at Moscow Zoo (Russia)


Chinese pygmy dormouse/ Typhlomys cinereus - ZooChat


Chinese Pigmy Dormouse Typhlomys cinereus
Three subspecies: cinereus, daloushanensis, jingdongensis. The subspecies chapensis from north Vietnam has been split as a full species, and the subspecies guangxiensis probably belongs in that species.


This species is not depicted here.

 

SPALACIDAE
Mole-Rats and Bamboo Rats
About thirty to forty species in six genera, of which only six species from four genera are represented in the Zoochat galleries.


This family of largely-fossorial rodents is divided into three or four subfamilies:

Myospalacinae (zokors) - about nine species in two genera (Eospalax, Myospalax)
None of the species are depicted here.

Rhizomyinae (bamboo rats and African mole-rats) - between six and seventeen species in three genera (Cannomys, Rhizomys, Tachyoryctes). Some authors would split this into two subfamilies: Rhizomyinae for the bamboo rats, and Tachyoryctidae for the African mole-rats.
There are four species depicted here, representing all three genera.

Spalacinae (blind mole-rats) - about fifteen-plus species in the single genus Spalax (although Nannospalax may also be separated)
There are only two species depicted here.


The accounts will follow the order of genera as given above (alpabetically within subfamily).

 

Myospalacinae (zokors)
About nine species in two genera (Eospalax, Myospalax). The two genera may be combined as Myospalax by some authors.

None of the species are represented in the Zoochat galleries.


Eospalax
About six species

Plateau Zokor Eospalax baileyi (otherwise treated as a subspecies of E. fontanierii)
Gansu Zokor Eospalax cansus (otherwise treated as a subspecies of E. fontanierii)
Chinese Zokor Eospalax fontanierii
Rothschild's Zokor Eospalax rothschildi
Reddish Zokor Eospalax rufescens
Smith's Zokor Eospalax smithii


Myospalax
Three species

False Zokor Myospalax aspalax
Siberian Zokor Myospalax myospalax
Transbaikal Zokor Myospalax psilurus

 

Rhizomyinae (bamboo rats and African mole-rats)
Between six and seventeen species in three genera (Cannomys, Rhizomys, Tachyoryctes).

There are only four species in the Zoochat galleries, but representing all three genera.

The Subfamily is further divided into two Tribes, Rhizomyini for the Asian bamboo rats and Tachyoryctini for the African mole-rats. Some taxonomists would treat these as two Subfamilies rather than Tribes. Although both groups evolved in Asia, the latter is now found solely in Africa.


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Cannomys
One species


Lesser Bamboo Rat Cannomys badius
Probably monotypic although a number of subspecies have been used in the past, including castaneus, minor, pater, plumbescens.

Photo by @vogelcommando in a private collection


Lesser bamboo rat - ZooChat


Photo by @Zooish at Singapore Zoo (Singapore)


Lesser Bamboo Rat (Cannomys badius) - ZooChat


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Rhizomys
Three species, two of which are represented in the Zoohat galleries.


Hoary Bamboo Rat Rhizomys pruinosus
About five subspecies: latouchei, pannosus, pruinosus, senex, umbriceps

Photo by @Little orca at the Hisense Science Discovery Center (China) (unknown subspecies)


Hoary Bamboo Rats（ Rhizomys pruinosus） - ZooChat


Chinese Bamboo Rat Rhizomys sinensis
About seven subspecies: davidi, neowardi, pediculus, reductus, sinensis, vestitus, wardi

There are a few photos in the Zoochat galleries labelled as being this species but they appear to actually show R. sumatrensis.
See Chinese bamboo rat (Rhizomys sinensis)? Species ID? - ZooChat and Chinese Bamboo Rat (Rhizomys sinensis) - ZooChat.


Large or Indomalayan Bamboo Rat Rhizomys sumatrensis
Four subspecies: cinereus, insularis, padangensis, sumatrensis

Photo by @Deer Forest at Chengdu Zanhuayuan Zoo (China) (subspecies cinereus)


large bamboo rat (Rhizomys sumatrensis) - ZooChat

Photo by @GregOz at Melaka Zoo (Malaysia) (subspecies sumatrensis)
(Despite the scientific name of this species, the type locality was actually Malacca [Melaka] in Peninsula Malaysia)


Bamboo Rat - Rhizomys sumatrensis - Melaka Zoo - 2009 - ZooChat

Photo by @Chlidonias at Melaka Zoo (Malaysia)
(The colouration does not seem to match that of the Malaysian subspecies, so it may be the Sumatran subspecies insularis for which the original description is "body silvery grey to yellowish", but it is probably more likely to just be individual variation)


Large bamboo rat (Rhizomys sumatrensis) - ZooChat

 

Tachyoryctes
Somewhere between two and thirteen species, only one of which is represented in the Zoochat galleries.

At its lowest limit the genus contains only two species, the Giant Mole-Rat T. macrocephalus and the East African Mole-Rat T. splendens. The opposite position is that T. splendens is a species-complex. Kingdon's Mammals of Africa presents just the two species, while the IUCN presents thirteen species.

The only species depicted in the Zoochat galleries is the Giant Mole-Rat.


Giant or Big-headed Mole-Rat Tachyoryctes macrocephalus
Two subspecies: hecki, macrocephalus

Photo by @Giant Eland in the wild, Ethiopia (subspecies hecki)


big-headed African mole rat, (Tachyoryctes macrocephalus), also known as the giant root-rat, Ethiopian African mole rat, or giant mole rat - ZooChat



The remaining species in the genus - if T. splendens is split up - are below:

Ankole Mole-Rat Tachyoryctes ankoliae
Mianzini Mole-Rat Tachyoryctes annectens
Aberdares Mole-Rat Tachyoryctes audax
Demon Mole-Rat Tachyoryctes daemon
Kenyan Mole-Rat Tachyoryctes ibeanus
Navivasha Mole-Rat Tachyoryctes naivashae
King Mole-Rat Tachyoryctes rex
Rwanda Mole-Rat Tachyoryctes ruandae
Rudd's Mole-Rat Tachyoryctes ruddi
Embi Mole-Rat Tachyoryctes spalacinus
Northeast African Mole-Rat Tachyoryctes splendens
Storey's Mole-Rat Tachyoryctes storeyi

 

Spalacinae (blind mole-rats)
About fifteen or more species in the single genus Spalax, although some small-bodied species may be separated as Nannospalax. Several of those listed below are probably species-complexes.

There are only two species represented in the Zoochat galleries. I have declined to list subspecies for these as taxonomic sources are extremely contradictory.



Palestine or Middle East Mole-Rat Spalax ehrenbergi
May also be placed in the separate genus Nannospalax.


Photo by @ralph at Pairi Daiza (Belgium)


Palestine Mole-rat - ZooChat


Upper Galilee Mountains Mole-Rat Spalax galili
May also be placed in the separate genus Nannospalax.

Photo by @Jakub of a captive animal at the University of South Bohemia (Czech Republic)


https://www.zoochat.com/community/m...ax-galili-university-of-south-bohemia.690524/



The remaining species in the genus - although not to be taken as any kind of definite list - are below:

Sandy Mole-Rat Spalax arenarius
Mount Carmel Mole-Rat Spalax carmeli
Giant Mole-Rat Spalax giganteus
Golan Heights Mole-Rat Spalax golani
Balkan Mole-Rat Spalax graecus
Judean Mountains Mole-Rat Spalax judaei
Lesser Mole-Rat Spalax leucodon
Greater Mole-Rat Spalax microphthalmus
Munzur Mole-Rat Spalax munzuri
Anatolian Mole-Rat Spalax nehringi
Kazakhstan Mole-Rat Spalax uralensis
Podolsk Mole-Rat Spalax zemni

 

Sadly no longer kept at Pairi Daiza - they were only present and onshow for about a year. Annoying, as I'm visiting in a little under a fortnight

 

CALOMYSCIDAE
Mouse-like Hamsters
Eight species in one genus, of which one species is represented in the Zoochat galleries.

Formerly included amongst the hamsters (in Cricetidae) due to similarities in the teeth and general appearance, although unlike true hamsters they have a long tail. All of the following species were previously combined as a single species, Calomyscus bailwardi.


Great Balkhan Mouse-like Hamster Calomyscus mystax
Monotypic

Photo by @vogelcommando at a private collection


Mouse-like hamster - ZooChat



The remaining species in this genus are as follows:

Zagros Mouse-like Hamster Calomyscus bailwardi
Baluchi Mouse-like Hamster Calomyscus baluchi
Goodwin's Mouse-like Hamster Calomyscus elburzensis
Noble Mouse-like Hamster Calomyscus grandis
Hotson's Mouse-like Hamster Calomyscus hotsoni
Tsolov's Mouse-like Hamster Calomyscus tsolovi
Urar Mouse-like Hamster Calomyscus urartensis